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The bond between Paladin and Lion was undefinable. Despite differences in species or cultures or opinions, it connected one person to four others with such depth that they could become one, avenging force of nature. It was the one thing that seemed to connect one small being to the whole universe and to fate itself. It was magical, strange, powerful, and a miracle.
And, unknown to the paladins, eternal.
(Or: what-if the paladins were able to communicate with the original paladins through their lions?)